The extensive debris that covers glaciers in the ablation zone of the Himalayan region plays an important part in regulating ablation rates and water availability for the downstream region. The melt rate of ice is determined by the amount of heat conducted through debris material lying over the ice. This study presents the vertical temperature gradients, thermal properties in terms of thermal diffusivity and thermal conductivity, and positive degree-day factors for the debris-covered portion of Lirung Glacier in Langtang Valley, Nepal Himalaya using field-based measurements from three different seasons. Field measurements include debris temperatures at different debris thicknesses, air temperature, and ice melt during the monsoon (2013), winter (2013), and pre-monsoon (2014) seasons. We used a thermal equation to estimate thermal diffusivity and thermal conductivity, and degree-day factors (DDF) were calculated from cumulative positive temperature and ice melt of the measurement period. Our analysis of debris temperature profiles at different depths of debris show the daily linear gradients of ?20.81 °C/m, 4.05 °C/m, and ?7.79 °C/m in the monsoon, winter, and pre-monsoon seasons, respectively. The values of thermal diffusivity and thermal conductivity in the monsoon season were 10 times greater than in the winter season. The large difference in these values is attributed to surface temperature and moisture content within the debris. Similarly, we found higher values of DDFs at thinner debris for the pre-monsoon season than in the monsoon season although we observed less melting during the pre-monsoon season. This is attributed to higher cumulative temperature during the monsoon season than in the pre-monsoon season. Our study advances our understanding of heat conductivity through debris material in different seasons, which supports estimating ice melt and discharge from glacierized river basins with debris-covered glaciers in the Himalayan region.

Understanding the variation in a plant's water sources is critical to understanding hydrological processes in water-limited environments. Here, we measured the stable-isotope ratios (δ¹⁸O) of xylem water of Caragana microphylla, precipitation, soil water from different depths, and groundwater to quantitatively analyze the proportion of water sources for the shrub. We found that the water sources of C. microphylla differed with the plant's ages and the seasons. The main water source for young shrubs was upper-soil water, and it showed significant changes with seasonal precipitation inputs. In summer, the proportion contributed by shallow water was significantly increased with increased precipitation inputs. Then, the contribution from shallow-soil water decreased with the decline in precipitation input in spring and autumn. However, the adult shrubs resorted to deep-soil layers and groundwater as the main water sources during the whole growing season and showed much less seasonal variation. We conclude that the main water source of the young shrubs was upper-soil water and was controlled by precipitation inputs. However, once the shrub gradually grew up and the roots reached sufficient depth, the main water sources change from the upper-soil layer recharged by precipitation to deep-soil water and groundwater, which were relatively stable and abundant in the desert ecosystem. These results also suggest that desert shrubs may be able to switch their main water sources to deep and reliable water sources as their age increases, and this adjustment to water availability carries significant importance for their acclimation to the desert habitat.

Soil calcium carbonate (CaCO₃) has a strong solid phosphorus effect, and high content of CaCO₃ can significantly reduce the effectiveness of soil phosphorus. To reveal the limiting effect of soil CaCO₃ on the growth of plants on sand land and its mechanism of plant physiology, we performed pot experiments with a two-factor randomized block design and a three-factor orthogonal design for different soil CaCO₃ content treatments using Artemisia ordosica seedlings. In the experiments, we surveyed plant height, aboveground biomass, root length and root weight and analyzed N, P concentrations and RNA content of the seedlings, and discussed the relationships between relative growth rate (RGR) of the seedlings and N:P ratio as well as RNA. Results show that, the RGRs of plant height and above-ground biomass of the seedlings decreased significantly with the increase of soil CaCO₃ content, and those for root length and root weight decreased. The RGRs of plant height and above-ground biomass of the seedlings were significantly negatively correlated with leaf N:P ratios, but significantly positively correlated with leaf RNA content and leaf P concentrations. It can be seen that soil CaCO₃ is a stress factor for the growth of A. ordosica seedlings, and the growth response of the seedlings under the influence of soil CaCO₃ is in line with the Growth Rate Hypothesis.

Although temperature extremes have led to more and more disasters, there are as yet few studies on the extremes and many disagreements on temperature changes in Antarctica. Based on daily minimum, maximum, and mean air temperatures (T_min, T_max, T_mean) at Great Wall Station (GW) and Zhongshan Station (ZS), we compared the temperature extremes and revealed a strong warming trend in T_min, a slight warming trend in T_mean, cooling in T_max, a decreasing trend in the daily temperature range, and the typical characteristic of coreless winter temperature. There are different seasonal variabilities, with the least in summer. The continentality index and seasonality show that the marine air mass has more effect on GW than ZS. Following the terminology of the Fifth Assessment Report of the Intergovernmental Panel on Climate Change (IPCC AR5), we defined nine indices of temperature extremes, based on the Antarctic geographical environment. Extreme-warm days have decreased, while extreme-warm nights have shown a nonsignificant trend. The number of melting days has increased at GW, while little change at ZS. More importantly, we have found inverse variations in temperature patterns between the two stations, which need further investigation into the dynamics of climate change in Antarctica.

Lakes regulate the water and heat exchange between the ground and the atmosphere on different temporal and spatial scales. However, studies of the lake effect in the high-altitude Tibetan Plateau (TP) rarely have been performed until recently, and little attention has been paid to modelling of frozen lakes. In this study, the Weather Research and Forecasting Model (WRF v. 3.6.1) is employed to conduct three numerical experiments in the Ngoring Lake Basin (the original experiment, an experiment with a tuned model, and a no-lake experiment) to investigate the influences of parameter optimization on the lake simulation and of the high-altitude lake on the regional climate. After the lake depth, the roughness lengths, and initial surface temperature are corrected in the model, the simulation of the air temperature is distinctly improved. In the experiment using a tuned model, the simulated sensible-heat flux (H) is clearly improved, especially during periods of ice melting (from late spring to early summer) and freezing (late fall). The improvement of latent-heat flux (LE) is mainly manifested by the sharp increase in the correlation coefficient between simulation and observation, whereas the improvement in the average value is small. The optimization of initial surface temperature shows the most prominent effect in the first year and distinctly weakens after a freezing period. After the lakes become grassland in the model, the daytime temperature clearly increases during the freezing and melting periods; but the nocturnal cooling appears in other stages, especially from September to October. The annual mean H increases by 6.4 times in the regions of the Ngoring Lake and the Gyaring Lake, and the LE declines by 56.2%. The sum of H and LE increases from 71.2 W/m² (with lake) to 84.6 W/m² (no lake). For the entire simulation region, the sum of H and LE also increases slightly. After the lakes are removed, the air temperature increases significantly from June to September over the area corresponding to the two lakes, and an abnormal convergence field appears; at the same time, the precipitation clearly increases over the two lakes and surrounding areas.

This article attempts to investigate the measure effect of rubble roadbed engineering in permafrost regions of Qinghai-Tibet Plateau. As a case study, Chaidaer-Muli Railway is used to evaluate the measure effect of rubble roadbed engineering in permafrost regions. The AHP (Analytic Hierarchy Process) method is thus employed to establish the evaluation indicator system. The evaluation factor is selected by analyzing the mutual relation between the permafrost environment and roadbed engineering. Thus, a hierarchical structure model is established based on the selected evaluation indices. Each factor is weighted to determine the status in the evaluation system, and grading standards are built for providing a basis for the evaluation. Then, the fuzzy mathematical method is introduced to evaluate the measure effect of rubble roadbed engineering in permafrost regions along the Chadaer-Muli Railway. Results show that most of the permafrost roadbed is in a preferable condition (b) along the Chaidaer-Muli Railway due to rubble engineering measures. This proportion reaches to 86.1%. The proportion in good (a), general (c) and poor states (d) are 0.0%, 7.5% and 6.4%, respectively, in all the evaluation sections along the Chaidaer-Muli Railway. Ground-temperature monitoring results are generally consistent with AHP-FUZZY evaluation results. This means that the AHP-FUZZY method can be applied to evaluate the effect of rubble roadbed engineering measures in permafrost regions. The effect evaluation of engineering measures will provide timely and effective feedback information for further engineering design. The series of engineering measures will more effectively protect permafrost stability.

For estimating the altitude-distribution pattern of carbon stocks in desert grasslands and analyzing the possible mechanism for this distribution, a detailed study was performed through a series of field vegetation surveys and soil samplings from 90 vegetation plots and 45 soil profiles at 9 sites of the Hexi Corridor region, Northwestern China. Aboveground, belowground, and litter-fall biomass-carbon stocks ranged from 43 to 109, 23 to 64, and 5 to 20 g/m², with mean values of 80.82, 44.91, and 12.15 g/m², respectively. Soil-carbon stocks varied between 2.88 and 3.98 kg/m², with a mean value of 3.43 kg/m² in the 0–100-cm soil layer. Both biomass- and soil-carbon stocks had an increasing tendency corresponding to the altitudinal gradient. A significantly negative correlation was found between soil-carbon stock and mean annual temperature, with further better correlations between soil- and biomass-carbon stocks, and mean annual precipitation. Furthermore, soil carbon was found to be positively correlated with soil-silt and -clay content, and negatively correlated with soil bulk density and the volume percent of gravel. It can be concluded that variations in soil texture and climate condition were the key factors influencing the altitudinal pattern of carbon stocks in this desert-grassland ecosystem. Thus, by using the linear-regression functions between altitude and carbon stocks, approximately 4.18 Tg carbon were predicted from the 1,260 km² of desert grasslands in the study area.

Iron deficiency chlorosis of Lilium davidii var. unicolor is often the case in practice in alkaline soils of northwest region of China. It is difficult to control iron chlorosis because of high cost and short effective work time of conventional iron fertilizers. In this study, a 2-year field experiment was conducted to evaluate the effects of two slow-release fertilizers on the suppression of iron deficiency chlorosis, soil chemical properties, and the yield and quality of L. davidii var. unicolor. Results show that both coated slow-release iron fertilizers and embedded slow-release iron fertilizer effectively controlled iron-deficiency chlorosis. The application of slow-release iron fertilizers significantly increased plant height and chlorophyll content of L. davidii var. unicolor at different growth stages. Furthermore, coated iron fertilizer application significantly increased starch, protein, soluble sugar and vitamin C content of L. davidii var. unicolor, and it also significantly improved total amino acid content, with increases in essential amino acids (Trp, Leu, Lys, Phe, Val, and Thr contents) and in nonessential amino acids (Asp, Glu, Cit, Ihs, Acc, Ala, Pro, and Cys contents). It was concluded that application of coated slow-release iron fertilizer could be a promising option for suppression of iron deficiency chlorosis and deserves further study.

Precipitation is an important component of global water and energy transport and a major aspect of climate change. Due to the scarcity of meteorological observations, the precipitation climate over Tibet has been insufficiently documented. In this study, the distribution of precipitation during the rainy season over Tibet from 1980 to 2013 is described on monthly to annual time scales with meteorological observations. Furthermore, four precipitation products are compared to observations over Tibet. These datasets include products derived from the Asian Precipitation-Highly-Resolved Observational Data (APHRO), the Global Precipitation Climatology Centre (GPCC), the University of Delaware (UDel), and the China Meteorological Administration (CMA). The error, relative error, standard deviation, root-mean-square error, correlations and trends between these products for the same period are analyzed with in situ precipitation during the rainy season from May to September. The results indicate that these datasets can broadly capture the temporal and spatial precipitation distribution over Tibet. The precipitation gradually increases from northwest to southeast. The spatial precipitation in GPCC and CMA are similar and positively correlated to observations. Areas with the largest deviations are located in southwestern Tibet along the Himalayas. The APHRO product underestimates, while the UDel, GPCC, and CMA datasets overestimates precipitation on the basis of monthly and inter-annual variation. The biases in GPCC and CMA are smaller than those in APHRO and UDel with a mean relative error lower than 10% during the same periods. The linear trend of precipitation indicates that the increase in precipitation has accelerated extensively during the last 30 years in most regions of Tibet. The CMA generally achieves the best performance of these four precipitation products. Data uncertainty in Tibet might be caused by the low density of stations, complex topography between the grid points and stations, and the interpolation methods, which can also produce an obvious difference between the gridded data and observations.

Most previous research on areas with abundant rainfall shows that simulations using rainfall-runoff modes have a very high prediction accuracy and applicability when using a back-propagation (BP), feed-forward, multilayer perceptron artificial neural network (ANN). However, in runoff areas with relatively low rainfall or a dry climate, more studies are needed. In these areas—of which oasis-plain areas are a particularly good example—the existence and development of runoff depends largely on that which is generated from alpine regions. Quantitative analysis of the uncertainty of runoff simulation under climate change is the key to improving the utilization and management of water resources in arid areas. Therefore, in this context, three kinds of BP feed-forward, three-layer ANNs with similar structure were chosen as models in this paper. Taking the oasis–plain region traverse by the Qira River Basin in Xinjiang, China, as the research area, the monthly accumulated runoff of the Qira River in the next month was simulated and predicted. The results showed that the training precision of a compact wavelet neural network is low; but from the forecasting results, it could be concluded that the training algorithm can better reflect the whole law of samples. The traditional artificial neural network (TANN) model and radial basis-function neural network (RBFNN) model showed higher accuracy in the training and prediction stage. However, the TANN model, more sensitive to the selection of input variables, requires a large number of numerical simulations to determine the appropriate input variables and the number of hidden-layer neurons. Hence, The RBFNN model is more suitable for the study of such problems. And it can be extended to other similar research arid-oasis areas on the southern edge of the Kunlun Mountains and provides a reference for sustainable water-resource management of arid-oasis areas.

Natural resource-management studies have become increasingly attentive to the influences of human factors. Among these, cultural biases shape people's responses to changes in natural resource systems. Several studies have applied grid-group cultural theory to assess the effects of multiple value biases among stakeholders on natural resource management. We developed and administered a questionnaire in the Heihe River Basin (n = 364) in northwestern China to investigate the appropriateness of applying this theory in the Chinese context of natural resource management. The results revealed various cultural biases among the respondents. In descending order of prevalence, these biases were hierarchism (46.98%), individualism (26.65%), egalitarianism (18.96%), and fatalism (2.78%), with the remaining respondents (4.67%) evidencing no obvious bias. Our empirical study revealed respondents' worldviews and the influence of sociodemographic characteristics on cultural biases, as theoretically posited. Among the variables examined, age had a positive and significant effect across all biases except individualism. The correlation of income to all cultural biases was consistently negative. Only education had a negative and significant effect across all biases. Women were found to adhere to egalitarianism, whereas men adhered to individualism and hierarchism. Thus, grid-group cultural theory was found to be appropriate in the Chinese context, with gender, age, education, and income evidently accounting for cultural biases. Relationships between environmental attitudes and cultural biases conformed with the hypothesis advanced by grid-group cultural theory. This finding may be of value in explaining individuals' environmental attitudes and facilitating the development and implementation of natural resource-management policies.

An automatic weather station (AWS) has been installed at the Qomolangma Station of the China Academy of Sciences (QOMS) since 2005, in a northern Himalayan valley near Mount Everest, with an altitude of 4,270 m a.s.l.. Nine years of meteorological records (2006–2014) from the automatic weather station (AWS) were analyzed in this study, aiming to understand the response of local weather to the seasonal transition on the northern slopes of Mount Everest, with consideration of the movement of the subtropical jet (STJ) and the onset of the Indian Summer Monsoon (ISM). We found: (1) Both the synoptic circulation and the orography have a profound influence on the local weather, especially the local circulation. (2) Southwesterly (SW) and southeasterly (SE) winds prevail alternately at QOMS in the afternoon through the year. The SW wind was driven by the STJ during the non-monsoon months, while the SE was induced by the trans-Himalayan flow through the Arun Valley, a major valley to the east of Mount Everest, under a background of weak westerly winds aloft. (3) The response of air temperature (T) and specific humidity (q) to the monsoon onset is not as marked as that of the nearsurface winds. The q increases gradually and reaches a maximum in July when the rainy period begins. (4) The alternation between the SW wind at QOMS and the afternoon SE wind in the pre-monsoon season signals the northward shift of the STJ and imminent monsoon onset. The average interval between these two events is 14 days.

Grain-size class-Std (GSCStd) and Grain-size class-dD (GSCdD) methods are simple statistical approaches for classifying bulk grain-size distributions (GSDs) into grain-size fractions. Although these two methods were developed based on similar statistical principles, the classification difference between these two methods has not been analyzed. In this study, GSCStd and GSCdD methods are conducted in thirteen grain-size data sequences to examine the applicability for identifying grain size fractions. Results show that, application of the GSCStd method is equivalent to that of the GSCdD method in identifying finer grain-size fractions, and the difference between the two methods mainly comes from the identification of coarse grain-size fractions. Thus, finer grain-size fractions are recommended for use in research of surface aeolian and paleo-aeolian sediments. In addition, our results do not completely agree with previous studies, coarser grain-size fractions in our case suggest that the GSCdD method may not be more applicable than the GSCStd method.

The Brassicaceae species Braya humilis shows broad adaptation to different climatic zones and latitudes. However, the molecular adaptation mechanism of B. humilis is poorly understood. In China, B. humilis is mainly distributed on the Qinghai-Tibetan Plateau (QTP) and in the adjacent arid region. Previous transcriptome analysis of B. humilis has revealed that 39 salt and osmotic stress response genes are subjected to purifying selection during its speciation. To further explore the adaptation mechanism of B. humilis to an arid environment, OrthoMCL program was employed in this study and 6,268 pairs of orthologous gene pairs with high confidence were obtained betweenB. humilis and Arabidopsis thaliana. A comparative evolutionary analysis based on nonsynonymous to synonymous substitution ratio (Ka/Ks) was then conducted. There were 64 pairs exhibiting a Ka/Ks ratio more than 0.5 and among which, three instrumental candidate genes, T2_20487, T2_22576, and T2_13757, were identified with strong selection signatures (Ka/Ks >1). The corresponding A. thaliana orthologs are double-stranded RNA-binding domain protein, MADS-box family protein, and NADH-dehydrogenase subunit 6, which is encoded by mitochondria genome. This report not only demonstrates the adaptation contribution of fast evolving nuclear genes, but also highlights the potential adaptive value of mitochondria gene to the speciation and adaptation of B. humilis toward the extreme environment in an arid region.

An underlying wetland surface comprises soil, water and vegetation and is sensitive to local climate change. Analysis of the degree of coupling between wetlands and the atmosphere and a quantitative assessment of how environmental factors influence latent heat flux have considerable scientific significance. Using data from observational tests of the Maduo Observatory of Climate and Environment of the Northwest Institute of Eco-Environment and Resource, CAS, from June 1 to August 31, 2014, this study analysed the time-varying characteristics and causes of the degree of coupling (Ω factor) between alpine wetlands underlying surface and the atmosphere and quantitatively calculated the influences of different environmental factors (solar radiation and vapour pressure deficit) on latent heat flux. The results were as follows: (1) Due to diurnal variations of solar radiation and wind speed, a trend developed where diurnal variations of the Ω factor were small in the morning and large in the evening. Due to the vegetation growing cycle, seasonal variations of the Ω factor present a reverse "U" trend. These trends are similar to the diurnal and seasonal variations of the absolute control exercised by solar radiation over latent heat flux. This conforms to the Omega Theory. (2) The values for average absolute atmospheric factor (surface factor or total) control exercised by solar radiation and water vapour pressure are 0.20 (0.02 or 0.22) and 0.005 (?0.07 or ?0.06) W/(m²·Pa), respectively. Generally speaking, solar radiation and water vapour pressure deficit exert opposite forces on latent heat flux. (3) At the underlying alpine wetland surface, solar radiation primarily influences latent heat flux through its direct effects (atmospheric factor controls). Water vapour pressure deficit primarily influences latent heat flux through its indirect effects (surface factor controls) on changing the surface resistance. (4) The average Ω factor in the underlying alpine wetland surface is high during the vegetation growing season, with a value of 0.38, and the degree of coupling between alpine wetland surface and atmosphere system is low. The actual measurements agree with the Omega Theory. The latent heat flux is mainly influenced by solar radiation.

Understanding the effect of biodiversity on ecosystem function is critical to promoting the sustainability of ecosystems and species conservation in natural ecosystems. We observed species composition, species richness and aboveground biomass, and simulated the competitive assemblages in a natural grassland ecosystem of China, aiming to test some assumptions and predictions about biodiversity–stability relationships. Our results show that aboveground productivity and temporal stability increased significantly with increasing species richness, and via a combination of overyielding, species asynchrony, and portfolio effects. Species interactions resulted in overyielding caused by trait-independent complementarity, and were not offset by a negative dominance effect and trait-dependent complementarity effect. Therefore, the mechanisms underlying the biodiversity effect shifted from the selection effect to the complementarity effect as diversity increased, and both effects were coexisted but the complementarity effect represent a mechanism that facilitates long term species coexistence in a natural grassland ecosystem of China.

To explore the characteristics of plant calcium (Ca) fraction, we analyzed 91 plant species in the Ningxia Habahu National Nature Reserve in Yanchi County of Ningxia and 84 plant species in Zhenglan Banner of Inner Mongolia. Results show that, for the two regions, there is no significant difference between Ca fraction for the same growth type, and between water soluble Ca content or between hydrochloric acid soluble Ca content for plants of the same ecosystem and between hydrochloric acid soluble Ca content for plants of the same family. In similar vegetation zones, there is a significant difference among hydrochloric acid soluble Ca content for different growth types in Yanchi County and Zhenglan Banner, which was the highest for annual herbs and the lowest for perennial herbs. There is a significant difference between acetic acid soluble Ca content and between hydrochloric acid soluble Ca content for sandy land and grassland ecosystems in Yanchi County. There is a significant difference among the same Ca fraction of different families in the same region. Thus, the characteristics of plant Ca fraction are results of long-term adaptation to the environment.

This work presents a reference system overview to improve the efficiency of biological improvement of saline-alkali soil developed during the last thirty years, ranging from connotation, general methods and species, soil desalination, soil structure, soil organic content, microbial flora, enzyme activity, yield and economic benefits. The reference system presented is divided into three main groups: suitable varieties, suitable cultivation measures, and a comprehensive evaluation system. There has been a lot of research on biological improvement of saline alkali soil, but these studies are very fragmented and lack a comprehensive standard system. Also, there is a lack of practical significance, particularly with regard to optimal species, densities and times of sowing for particular regions. On the other hand, the corresponding cultivation measure is very important. Therefore, a reference system plays an important role to the effect of biological improvement of saline alkali soil.

Mu Us Sandy Land (MUSL) of China, as a typical eco-fragile and farming-pastoral transitional region, shows great vulnerability to disturbances from cultivation activity. In this region, the conflict between cultivation activity and environmental protection has not attracted great importance until the implementation of China's Grain-for-Green Policy (CGGP) since 2000. Here, using Landsat5 TM/Landsat7 ETM+ images from 1990, 2000, and 2010, we monitor the cultivation activity and land-use/cover changes (LUCC) resulting from cultivation activity in the MUSL region. Based on the data from images, we developed a series of databases of cultivated activity-induced LUCC and use them to discuss comparatively the spatio-temporal evolution trends of cultivation activity before and after CGGP implementation. These results provide evidence for managers to evaluate the implementation effectiveness of governmental policy and the influence of cultivation activity on the ecological environment of the MUSL region.

An overall greening over the Tibetan Plateau (TP) in recent decades has been established through analyses of remotely sensed Normalized Difference Vegetation Index (NDVI), though the regional pattern of the changes and associated drivers remain to be explored. This study used a satellite Leaf Area Index (LAI) dataset (the GLASS LAI dataset) and examined vegetation changes in humid and arid regions of the TP during 1982–2012. Based on distributions of the major vegetation types, the TP was divided roughly into a humid southeastern region dominated by meadow and a dry northwestern region covered mainly by steppe. It was found that the dividing line between the two regions corresponded well with the lines of mean annual precipitation of 400 mm and the mean LAI of 0.3. LAI=0.3 was subsequently used as a threshold for investigating vegetation type changes at the interanual and decadal time scales: if LAI increased from less than 0.3 to greater than 0.3 from one time period to the next, it was regarded as a change from steppe to meadow, and vice versa. The analysis shows that changes in vegetation types occurred primarily around the dividing line of the two regions, with clear growth (reduction) of the area covered by meadow (steppe), in consistency with the findings from using another independent satellite product. Surface air temperature and precipitation (diurnal temperature range) appeared to contribute positively (negatively) to this change though climate variables displayed varying correlation with LAI for different time periods and different regions.

Khaya belongs to the family Meliaceae. In Nigeria the genus is represented by three species viz; K. senegalensis A. Juss., K. grandifoliola C. DC. and K. ivorensis A. Chev. Comparative foliar anatomy of the three Khaya species was carried out to identify and describe distinctive anatomical characters that could possibly be used to delimit the three taxa. Transverse section, epidermal peels and cleared leaves of these three species were made. Characteristic similarity and disparity in the tissues arrangement as well as cell inclusions were noted for description and delimitation. The three Khaya species studied had essentially the same anatomical features, e.g., venation pattern having open polygonal areoles and the veins terminals biforkated. However, there were characters that seem to be species specific, e.g., vien termination number and areole width. The leaf epidermal studies of the three species revealed similarities in stomatal type which are generally staurocytic, epidermal cells and undulating anticlinal cell walls but stomata density varied. Hexacytic stomata is only observed in the abaxial surface of K. grandifoliola which distinguished this species from the others. The leaf petiole shape of the three species are round and difficult to distinguish into adaxial and abaxial surfaces. The cuticle is striated, vascular bundles are heart shape, conjoint, concentric and amphivasal, but are different in epidermal and collenchyma cell layer numbers. The leaf transverse sections of the three Khaya species studied have conjoint, concentric and amphicribral bundles while the leaf cuticle of K. senegalensis and K. grandifoliola are striated but that of Khaya ivorensis is non-striated.

Desert plants take on unique physiologically adaptive mechanisms in response to an adverse environment. In this study, we determined the concentrations of leaf nitrogen (N), phosphorus (P), and calcium (Ca) fraction for dominant species of Artemisia ordosica, A. frigida, Calligonum mongolicum, and Oxytropis aciphylla in the Alxa Desert and discussed seasonal changes of their leaf N:P ratio and Ca fraction. The results showed that, from May to September, the N:P ratios of A. ordosica and C. mongolicum gradually and significantly increased, while those of A. frigida, and O. aciphylla had an increase trend that was not significant; the physiologically active Ca of A. ordosica and A. frigida increased significantly, while that of C. mongolicum and O. aciphylla decreased significantly. The physiologically inert calcium of C. mongolicum increased extremely significantly, while that of others was not significant. There was a significantly positive correlation between the N:P ratio and physiologically active Ca for A. ordosica, and the N:P ratio was significantly and negatively correlated with physiologically active Ca for O. aciphylla. These findings revealed that the physiological regulation mechanism was different for the plants either in earlier stage or later stage of plant-community succession.

The extensive debris that covers glaciers in the ablation zone of the Himalayan region plays an important part in regulating ablation rates and water availability for the downstream region. The melt rate of ice is determined by the amount of heat conducted through debris material lying over the ice. This study presents the vertical temperature gradients, thermal properties in terms of thermal diffusivity and thermal conductivity, and positive degree-day factors for the debris-covered portion of Lirung Glacier in Langtang Valley, Nepal Himalaya using field-based measurements from three different seasons. Field measurements include debris temperatures at different debris thicknesses, air temperature, and ice melt during the monsoon (2013), winter (2013), and pre-monsoon (2014) seasons. We used a thermal equation to estimate thermal diffusivity and thermal conductivity, and degree-day factors (DDF) were calculated from cumulative positive temperature and ice melt of the measurement period. Our analysis of debris temperature profiles at different depths of debris show the daily linear gradients of ?20.81 °C/m, 4.05 °C/m, and ?7.79 °C/m in the monsoon, winter, and pre-monsoon seasons, respectively. The values of thermal diffusivity and thermal conductivity in the monsoon season were 10 times greater than in the winter season. The large difference in these values is attributed to surface temperature and moisture content within the debris. Similarly, we found higher values of DDFs at thinner debris for the pre-monsoon season than in the monsoon season although we observed less melting during the pre-monsoon season. This is attributed to higher cumulative temperature during the monsoon season than in the pre-monsoon season. Our study advances our understanding of heat conductivity through debris material in different seasons, which supports estimating ice melt and discharge from glacierized river basins with debris-covered glaciers in the Himalayan region.

Understanding the variation in a plant's water sources is critical to understanding hydrological processes in water-limited environments. Here, we measured the stable-isotope ratios (δ¹⁸O) of xylem water of Caragana microphylla, precipitation, soil water from different depths, and groundwater to quantitatively analyze the proportion of water sources for the shrub. We found that the water sources of C. microphylla differed with the plant's ages and the seasons. The main water source for young shrubs was upper-soil water, and it showed significant changes with seasonal precipitation inputs. In summer, the proportion contributed by shallow water was significantly increased with increased precipitation inputs. Then, the contribution from shallow-soil water decreased with the decline in precipitation input in spring and autumn. However, the adult shrubs resorted to deep-soil layers and groundwater as the main water sources during the whole growing season and showed much less seasonal variation. We conclude that the main water source of the young shrubs was upper-soil water and was controlled by precipitation inputs. However, once the shrub gradually grew up and the roots reached sufficient depth, the main water sources change from the upper-soil layer recharged by precipitation to deep-soil water and groundwater, which were relatively stable and abundant in the desert ecosystem. These results also suggest that desert shrubs may be able to switch their main water sources to deep and reliable water sources as their age increases, and this adjustment to water availability carries significant importance for their acclimation to the desert habitat.

Soil calcium carbonate (CaCO₃) has a strong solid phosphorus effect, and high content of CaCO₃ can significantly reduce the effectiveness of soil phosphorus. To reveal the limiting effect of soil CaCO₃ on the growth of plants on sand land and its mechanism of plant physiology, we performed pot experiments with a two-factor randomized block design and a three-factor orthogonal design for different soil CaCO₃ content treatments using Artemisia ordosica seedlings. In the experiments, we surveyed plant height, aboveground biomass, root length and root weight and analyzed N, P concentrations and RNA content of the seedlings, and discussed the relationships between relative growth rate (RGR) of the seedlings and N:P ratio as well as RNA. Results show that, the RGRs of plant height and above-ground biomass of the seedlings decreased significantly with the increase of soil CaCO₃ content, and those for root length and root weight decreased. The RGRs of plant height and above-ground biomass of the seedlings were significantly negatively correlated with leaf N:P ratios, but significantly positively correlated with leaf RNA content and leaf P concentrations. It can be seen that soil CaCO₃ is a stress factor for the growth of A. ordosica seedlings, and the growth response of the seedlings under the influence of soil CaCO₃ is in line with the Growth Rate Hypothesis.

Although temperature extremes have led to more and more disasters, there are as yet few studies on the extremes and many disagreements on temperature changes in Antarctica. Based on daily minimum, maximum, and mean air temperatures (T_min, T_max, T_mean) at Great Wall Station (GW) and Zhongshan Station (ZS), we compared the temperature extremes and revealed a strong warming trend in T_min, a slight warming trend in T_mean, cooling in T_max, a decreasing trend in the daily temperature range, and the typical characteristic of coreless winter temperature. There are different seasonal variabilities, with the least in summer. The continentality index and seasonality show that the marine air mass has more effect on GW than ZS. Following the terminology of the Fifth Assessment Report of the Intergovernmental Panel on Climate Change (IPCC AR5), we defined nine indices of temperature extremes, based on the Antarctic geographical environment. Extreme-warm days have decreased, while extreme-warm nights have shown a nonsignificant trend. The number of melting days has increased at GW, while little change at ZS. More importantly, we have found inverse variations in temperature patterns between the two stations, which need further investigation into the dynamics of climate change in Antarctica.

Lakes regulate the water and heat exchange between the ground and the atmosphere on different temporal and spatial scales. However, studies of the lake effect in the high-altitude Tibetan Plateau (TP) rarely have been performed until recently, and little attention has been paid to modelling of frozen lakes. In this study, the Weather Research and Forecasting Model (WRF v. 3.6.1) is employed to conduct three numerical experiments in the Ngoring Lake Basin (the original experiment, an experiment with a tuned model, and a no-lake experiment) to investigate the influences of parameter optimization on the lake simulation and of the high-altitude lake on the regional climate. After the lake depth, the roughness lengths, and initial surface temperature are corrected in the model, the simulation of the air temperature is distinctly improved. In the experiment using a tuned model, the simulated sensible-heat flux (H) is clearly improved, especially during periods of ice melting (from late spring to early summer) and freezing (late fall). The improvement of latent-heat flux (LE) is mainly manifested by the sharp increase in the correlation coefficient between simulation and observation, whereas the improvement in the average value is small. The optimization of initial surface temperature shows the most prominent effect in the first year and distinctly weakens after a freezing period. After the lakes become grassland in the model, the daytime temperature clearly increases during the freezing and melting periods; but the nocturnal cooling appears in other stages, especially from September to October. The annual mean H increases by 6.4 times in the regions of the Ngoring Lake and the Gyaring Lake, and the LE declines by 56.2%. The sum of H and LE increases from 71.2 W/m² (with lake) to 84.6 W/m² (no lake). For the entire simulation region, the sum of H and LE also increases slightly. After the lakes are removed, the air temperature increases significantly from June to September over the area corresponding to the two lakes, and an abnormal convergence field appears; at the same time, the precipitation clearly increases over the two lakes and surrounding areas.

This article attempts to investigate the measure effect of rubble roadbed engineering in permafrost regions of Qinghai-Tibet Plateau. As a case study, Chaidaer-Muli Railway is used to evaluate the measure effect of rubble roadbed engineering in permafrost regions. The AHP (Analytic Hierarchy Process) method is thus employed to establish the evaluation indicator system. The evaluation factor is selected by analyzing the mutual relation between the permafrost environment and roadbed engineering. Thus, a hierarchical structure model is established based on the selected evaluation indices. Each factor is weighted to determine the status in the evaluation system, and grading standards are built for providing a basis for the evaluation. Then, the fuzzy mathematical method is introduced to evaluate the measure effect of rubble roadbed engineering in permafrost regions along the Chadaer-Muli Railway. Results show that most of the permafrost roadbed is in a preferable condition (b) along the Chaidaer-Muli Railway due to rubble engineering measures. This proportion reaches to 86.1%. The proportion in good (a), general (c) and poor states (d) are 0.0%, 7.5% and 6.4%, respectively, in all the evaluation sections along the Chaidaer-Muli Railway. Ground-temperature monitoring results are generally consistent with AHP-FUZZY evaluation results. This means that the AHP-FUZZY method can be applied to evaluate the effect of rubble roadbed engineering measures in permafrost regions. The effect evaluation of engineering measures will provide timely and effective feedback information for further engineering design. The series of engineering measures will more effectively protect permafrost stability.

For estimating the altitude-distribution pattern of carbon stocks in desert grasslands and analyzing the possible mechanism for this distribution, a detailed study was performed through a series of field vegetation surveys and soil samplings from 90 vegetation plots and 45 soil profiles at 9 sites of the Hexi Corridor region, Northwestern China. Aboveground, belowground, and litter-fall biomass-carbon stocks ranged from 43 to 109, 23 to 64, and 5 to 20 g/m², with mean values of 80.82, 44.91, and 12.15 g/m², respectively. Soil-carbon stocks varied between 2.88 and 3.98 kg/m², with a mean value of 3.43 kg/m² in the 0–100-cm soil layer. Both biomass- and soil-carbon stocks had an increasing tendency corresponding to the altitudinal gradient. A significantly negative correlation was found between soil-carbon stock and mean annual temperature, with further better correlations between soil- and biomass-carbon stocks, and mean annual precipitation. Furthermore, soil carbon was found to be positively correlated with soil-silt and -clay content, and negatively correlated with soil bulk density and the volume percent of gravel. It can be concluded that variations in soil texture and climate condition were the key factors influencing the altitudinal pattern of carbon stocks in this desert-grassland ecosystem. Thus, by using the linear-regression functions between altitude and carbon stocks, approximately 4.18 Tg carbon were predicted from the 1,260 km² of desert grasslands in the study area.

Iron deficiency chlorosis of Lilium davidii var. unicolor is often the case in practice in alkaline soils of northwest region of China. It is difficult to control iron chlorosis because of high cost and short effective work time of conventional iron fertilizers. In this study, a 2-year field experiment was conducted to evaluate the effects of two slow-release fertilizers on the suppression of iron deficiency chlorosis, soil chemical properties, and the yield and quality of L. davidii var. unicolor. Results show that both coated slow-release iron fertilizers and embedded slow-release iron fertilizer effectively controlled iron-deficiency chlorosis. The application of slow-release iron fertilizers significantly increased plant height and chlorophyll content of L. davidii var. unicolor at different growth stages. Furthermore, coated iron fertilizer application significantly increased starch, protein, soluble sugar and vitamin C content of L. davidii var. unicolor, and it also significantly improved total amino acid content, with increases in essential amino acids (Trp, Leu, Lys, Phe, Val, and Thr contents) and in nonessential amino acids (Asp, Glu, Cit, Ihs, Acc, Ala, Pro, and Cys contents). It was concluded that application of coated slow-release iron fertilizer could be a promising option for suppression of iron deficiency chlorosis and deserves further study.

Precipitation is an important component of global water and energy transport and a major aspect of climate change. Due to the scarcity of meteorological observations, the precipitation climate over Tibet has been insufficiently documented. In this study, the distribution of precipitation during the rainy season over Tibet from 1980 to 2013 is described on monthly to annual time scales with meteorological observations. Furthermore, four precipitation products are compared to observations over Tibet. These datasets include products derived from the Asian Precipitation-Highly-Resolved Observational Data (APHRO), the Global Precipitation Climatology Centre (GPCC), the University of Delaware (UDel), and the China Meteorological Administration (CMA). The error, relative error, standard deviation, root-mean-square error, correlations and trends between these products for the same period are analyzed with in situ precipitation during the rainy season from May to September. The results indicate that these datasets can broadly capture the temporal and spatial precipitation distribution over Tibet. The precipitation gradually increases from northwest to southeast. The spatial precipitation in GPCC and CMA are similar and positively correlated to observations. Areas with the largest deviations are located in southwestern Tibet along the Himalayas. The APHRO product underestimates, while the UDel, GPCC, and CMA datasets overestimates precipitation on the basis of monthly and inter-annual variation. The biases in GPCC and CMA are smaller than those in APHRO and UDel with a mean relative error lower than 10% during the same periods. The linear trend of precipitation indicates that the increase in precipitation has accelerated extensively during the last 30 years in most regions of Tibet. The CMA generally achieves the best performance of these four precipitation products. Data uncertainty in Tibet might be caused by the low density of stations, complex topography between the grid points and stations, and the interpolation methods, which can also produce an obvious difference between the gridded data and observations.

Most previous research on areas with abundant rainfall shows that simulations using rainfall-runoff modes have a very high prediction accuracy and applicability when using a back-propagation (BP), feed-forward, multilayer perceptron artificial neural network (ANN). However, in runoff areas with relatively low rainfall or a dry climate, more studies are needed. In these areas—of which oasis-plain areas are a particularly good example—the existence and development of runoff depends largely on that which is generated from alpine regions. Quantitative analysis of the uncertainty of runoff simulation under climate change is the key to improving the utilization and management of water resources in arid areas. Therefore, in this context, three kinds of BP feed-forward, three-layer ANNs with similar structure were chosen as models in this paper. Taking the oasis–plain region traverse by the Qira River Basin in Xinjiang, China, as the research area, the monthly accumulated runoff of the Qira River in the next month was simulated and predicted. The results showed that the training precision of a compact wavelet neural network is low; but from the forecasting results, it could be concluded that the training algorithm can better reflect the whole law of samples. The traditional artificial neural network (TANN) model and radial basis-function neural network (RBFNN) model showed higher accuracy in the training and prediction stage. However, the TANN model, more sensitive to the selection of input variables, requires a large number of numerical simulations to determine the appropriate input variables and the number of hidden-layer neurons. Hence, The RBFNN model is more suitable for the study of such problems. And it can be extended to other similar research arid-oasis areas on the southern edge of the Kunlun Mountains and provides a reference for sustainable water-resource management of arid-oasis areas.

Natural resource-management studies have become increasingly attentive to the influences of human factors. Among these, cultural biases shape people's responses to changes in natural resource systems. Several studies have applied grid-group cultural theory to assess the effects of multiple value biases among stakeholders on natural resource management. We developed and administered a questionnaire in the Heihe River Basin (n = 364) in northwestern China to investigate the appropriateness of applying this theory in the Chinese context of natural resource management. The results revealed various cultural biases among the respondents. In descending order of prevalence, these biases were hierarchism (46.98%), individualism (26.65%), egalitarianism (18.96%), and fatalism (2.78%), with the remaining respondents (4.67%) evidencing no obvious bias. Our empirical study revealed respondents' worldviews and the influence of sociodemographic characteristics on cultural biases, as theoretically posited. Among the variables examined, age had a positive and significant effect across all biases except individualism. The correlation of income to all cultural biases was consistently negative. Only education had a negative and significant effect across all biases. Women were found to adhere to egalitarianism, whereas men adhered to individualism and hierarchism. Thus, grid-group cultural theory was found to be appropriate in the Chinese context, with gender, age, education, and income evidently accounting for cultural biases. Relationships between environmental attitudes and cultural biases conformed with the hypothesis advanced by grid-group cultural theory. This finding may be of value in explaining individuals' environmental attitudes and facilitating the development and implementation of natural resource-management policies.

An automatic weather station (AWS) has been installed at the Qomolangma Station of the China Academy of Sciences (QOMS) since 2005, in a northern Himalayan valley near Mount Everest, with an altitude of 4,270 m a.s.l.. Nine years of meteorological records (2006–2014) from the automatic weather station (AWS) were analyzed in this study, aiming to understand the response of local weather to the seasonal transition on the northern slopes of Mount Everest, with consideration of the movement of the subtropical jet (STJ) and the onset of the Indian Summer Monsoon (ISM). We found: (1) Both the synoptic circulation and the orography have a profound influence on the local weather, especially the local circulation. (2) Southwesterly (SW) and southeasterly (SE) winds prevail alternately at QOMS in the afternoon through the year. The SW wind was driven by the STJ during the non-monsoon months, while the SE was induced by the trans-Himalayan flow through the Arun Valley, a major valley to the east of Mount Everest, under a background of weak westerly winds aloft. (3) The response of air temperature (T) and specific humidity (q) to the monsoon onset is not as marked as that of the nearsurface winds. The q increases gradually and reaches a maximum in July when the rainy period begins. (4) The alternation between the SW wind at QOMS and the afternoon SE wind in the pre-monsoon season signals the northward shift of the STJ and imminent monsoon onset. The average interval between these two events is 14 days.

Grain-size class-Std (GSCStd) and Grain-size class-dD (GSCdD) methods are simple statistical approaches for classifying bulk grain-size distributions (GSDs) into grain-size fractions. Although these two methods were developed based on similar statistical principles, the classification difference between these two methods has not been analyzed. In this study, GSCStd and GSCdD methods are conducted in thirteen grain-size data sequences to examine the applicability for identifying grain size fractions. Results show that, application of the GSCStd method is equivalent to that of the GSCdD method in identifying finer grain-size fractions, and the difference between the two methods mainly comes from the identification of coarse grain-size fractions. Thus, finer grain-size fractions are recommended for use in research of surface aeolian and paleo-aeolian sediments. In addition, our results do not completely agree with previous studies, coarser grain-size fractions in our case suggest that the GSCdD method may not be more applicable than the GSCStd method.

The Brassicaceae species Braya humilis shows broad adaptation to different climatic zones and latitudes. However, the molecular adaptation mechanism of B. humilis is poorly understood. In China, B. humilis is mainly distributed on the Qinghai-Tibetan Plateau (QTP) and in the adjacent arid region. Previous transcriptome analysis of B. humilis has revealed that 39 salt and osmotic stress response genes are subjected to purifying selection during its speciation. To further explore the adaptation mechanism of B. humilis to an arid environment, OrthoMCL program was employed in this study and 6,268 pairs of orthologous gene pairs with high confidence were obtained betweenB. humilis and Arabidopsis thaliana. A comparative evolutionary analysis based on nonsynonymous to synonymous substitution ratio (Ka/Ks) was then conducted. There were 64 pairs exhibiting a Ka/Ks ratio more than 0.5 and among which, three instrumental candidate genes, T2_20487, T2_22576, and T2_13757, were identified with strong selection signatures (Ka/Ks >1). The corresponding A. thaliana orthologs are double-stranded RNA-binding domain protein, MADS-box family protein, and NADH-dehydrogenase subunit 6, which is encoded by mitochondria genome. This report not only demonstrates the adaptation contribution of fast evolving nuclear genes, but also highlights the potential adaptive value of mitochondria gene to the speciation and adaptation of B. humilis toward the extreme environment in an arid region.

An underlying wetland surface comprises soil, water and vegetation and is sensitive to local climate change. Analysis of the degree of coupling between wetlands and the atmosphere and a quantitative assessment of how environmental factors influence latent heat flux have considerable scientific significance. Using data from observational tests of the Maduo Observatory of Climate and Environment of the Northwest Institute of Eco-Environment and Resource, CAS, from June 1 to August 31, 2014, this study analysed the time-varying characteristics and causes of the degree of coupling (Ω factor) between alpine wetlands underlying surface and the atmosphere and quantitatively calculated the influences of different environmental factors (solar radiation and vapour pressure deficit) on latent heat flux. The results were as follows: (1) Due to diurnal variations of solar radiation and wind speed, a trend developed where diurnal variations of the Ω factor were small in the morning and large in the evening. Due to the vegetation growing cycle, seasonal variations of the Ω factor present a reverse "U" trend. These trends are similar to the diurnal and seasonal variations of the absolute control exercised by solar radiation over latent heat flux. This conforms to the Omega Theory. (2) The values for average absolute atmospheric factor (surface factor or total) control exercised by solar radiation and water vapour pressure are 0.20 (0.02 or 0.22) and 0.005 (?0.07 or ?0.06) W/(m²·Pa), respectively. Generally speaking, solar radiation and water vapour pressure deficit exert opposite forces on latent heat flux. (3) At the underlying alpine wetland surface, solar radiation primarily influences latent heat flux through its direct effects (atmospheric factor controls). Water vapour pressure deficit primarily influences latent heat flux through its indirect effects (surface factor controls) on changing the surface resistance. (4) The average Ω factor in the underlying alpine wetland surface is high during the vegetation growing season, with a value of 0.38, and the degree of coupling between alpine wetland surface and atmosphere system is low. The actual measurements agree with the Omega Theory. The latent heat flux is mainly influenced by solar radiation.

Understanding the effect of biodiversity on ecosystem function is critical to promoting the sustainability of ecosystems and species conservation in natural ecosystems. We observed species composition, species richness and aboveground biomass, and simulated the competitive assemblages in a natural grassland ecosystem of China, aiming to test some assumptions and predictions about biodiversity–stability relationships. Our results show that aboveground productivity and temporal stability increased significantly with increasing species richness, and via a combination of overyielding, species asynchrony, and portfolio effects. Species interactions resulted in overyielding caused by trait-independent complementarity, and were not offset by a negative dominance effect and trait-dependent complementarity effect. Therefore, the mechanisms underlying the biodiversity effect shifted from the selection effect to the complementarity effect as diversity increased, and both effects were coexisted but the complementarity effect represent a mechanism that facilitates long term species coexistence in a natural grassland ecosystem of China.

To explore the characteristics of plant calcium (Ca) fraction, we analyzed 91 plant species in the Ningxia Habahu National Nature Reserve in Yanchi County of Ningxia and 84 plant species in Zhenglan Banner of Inner Mongolia. Results show that, for the two regions, there is no significant difference between Ca fraction for the same growth type, and between water soluble Ca content or between hydrochloric acid soluble Ca content for plants of the same ecosystem and between hydrochloric acid soluble Ca content for plants of the same family. In similar vegetation zones, there is a significant difference among hydrochloric acid soluble Ca content for different growth types in Yanchi County and Zhenglan Banner, which was the highest for annual herbs and the lowest for perennial herbs. There is a significant difference between acetic acid soluble Ca content and between hydrochloric acid soluble Ca content for sandy land and grassland ecosystems in Yanchi County. There is a significant difference among the same Ca fraction of different families in the same region. Thus, the characteristics of plant Ca fraction are results of long-term adaptation to the environment.

This work presents a reference system overview to improve the efficiency of biological improvement of saline-alkali soil developed during the last thirty years, ranging from connotation, general methods and species, soil desalination, soil structure, soil organic content, microbial flora, enzyme activity, yield and economic benefits. The reference system presented is divided into three main groups: suitable varieties, suitable cultivation measures, and a comprehensive evaluation system. There has been a lot of research on biological improvement of saline alkali soil, but these studies are very fragmented and lack a comprehensive standard system. Also, there is a lack of practical significance, particularly with regard to optimal species, densities and times of sowing for particular regions. On the other hand, the corresponding cultivation measure is very important. Therefore, a reference system plays an important role to the effect of biological improvement of saline alkali soil.

Mu Us Sandy Land (MUSL) of China, as a typical eco-fragile and farming-pastoral transitional region, shows great vulnerability to disturbances from cultivation activity. In this region, the conflict between cultivation activity and environmental protection has not attracted great importance until the implementation of China's Grain-for-Green Policy (CGGP) since 2000. Here, using Landsat5 TM/Landsat7 ETM+ images from 1990, 2000, and 2010, we monitor the cultivation activity and land-use/cover changes (LUCC) resulting from cultivation activity in the MUSL region. Based on the data from images, we developed a series of databases of cultivated activity-induced LUCC and use them to discuss comparatively the spatio-temporal evolution trends of cultivation activity before and after CGGP implementation. These results provide evidence for managers to evaluate the implementation effectiveness of governmental policy and the influence of cultivation activity on the ecological environment of the MUSL region.

An overall greening over the Tibetan Plateau (TP) in recent decades has been established through analyses of remotely sensed Normalized Difference Vegetation Index (NDVI), though the regional pattern of the changes and associated drivers remain to be explored. This study used a satellite Leaf Area Index (LAI) dataset (the GLASS LAI dataset) and examined vegetation changes in humid and arid regions of the TP during 1982–2012. Based on distributions of the major vegetation types, the TP was divided roughly into a humid southeastern region dominated by meadow and a dry northwestern region covered mainly by steppe. It was found that the dividing line between the two regions corresponded well with the lines of mean annual precipitation of 400 mm and the mean LAI of 0.3. LAI=0.3 was subsequently used as a threshold for investigating vegetation type changes at the interanual and decadal time scales: if LAI increased from less than 0.3 to greater than 0.3 from one time period to the next, it was regarded as a change from steppe to meadow, and vice versa. The analysis shows that changes in vegetation types occurred primarily around the dividing line of the two regions, with clear growth (reduction) of the area covered by meadow (steppe), in consistency with the findings from using another independent satellite product. Surface air temperature and precipitation (diurnal temperature range) appeared to contribute positively (negatively) to this change though climate variables displayed varying correlation with LAI for different time periods and different regions.

Khaya belongs to the family Meliaceae. In Nigeria the genus is represented by three species viz; K. senegalensis A. Juss., K. grandifoliola C. DC. and K. ivorensis A. Chev. Comparative foliar anatomy of the three Khaya species was carried out to identify and describe distinctive anatomical characters that could possibly be used to delimit the three taxa. Transverse section, epidermal peels and cleared leaves of these three species were made. Characteristic similarity and disparity in the tissues arrangement as well as cell inclusions were noted for description and delimitation. The three Khaya species studied had essentially the same anatomical features, e.g., venation pattern having open polygonal areoles and the veins terminals biforkated. However, there were characters that seem to be species specific, e.g., vien termination number and areole width. The leaf epidermal studies of the three species revealed similarities in stomatal type which are generally staurocytic, epidermal cells and undulating anticlinal cell walls but stomata density varied. Hexacytic stomata is only observed in the abaxial surface of K. grandifoliola which distinguished this species from the others. The leaf petiole shape of the three species are round and difficult to distinguish into adaxial and abaxial surfaces. The cuticle is striated, vascular bundles are heart shape, conjoint, concentric and amphivasal, but are different in epidermal and collenchyma cell layer numbers. The leaf transverse sections of the three Khaya species studied have conjoint, concentric and amphicribral bundles while the leaf cuticle of K. senegalensis and K. grandifoliola are striated but that of Khaya ivorensis is non-striated.

Desert plants take on unique physiologically adaptive mechanisms in response to an adverse environment. In this study, we determined the concentrations of leaf nitrogen (N), phosphorus (P), and calcium (Ca) fraction for dominant species of Artemisia ordosica, A. frigida, Calligonum mongolicum, and Oxytropis aciphylla in the Alxa Desert and discussed seasonal changes of their leaf N:P ratio and Ca fraction. The results showed that, from May to September, the N:P ratios of A. ordosica and C. mongolicum gradually and significantly increased, while those of A. frigida, and O. aciphylla had an increase trend that was not significant; the physiologically active Ca of A. ordosica and A. frigida increased significantly, while that of C. mongolicum and O. aciphylla decreased significantly. The physiologically inert calcium of C. mongolicum increased extremely significantly, while that of others was not significant. There was a significantly positive correlation between the N:P ratio and physiologically active Ca for A. ordosica, and the N:P ratio was significantly and negatively correlated with physiologically active Ca for O. aciphylla. These findings revealed that the physiological regulation mechanism was different for the plants either in earlier stage or later stage of plant-community succession.